Which preposition to use with each

OF Occurrences 1%

NEAR THIS SPOT WERE INTERRED THE MORTAL REMAINS OF ETHELBALD AND ETHELBERT HIS BROTHER EACH OF WHOM IN TURN SUCCEEDED TO THE THRONE OF ETHELWOULF THIER FATHER KING OF THE WEST SAXONS AND WERE SUCCEEDED IN THE KINGDOM

Which preposition to use with  each