Which preposition to use with hackles

of Occurrences 3%

The general colour of the former is golden, or orange-yellow, each feather having a glossy dark brown or black tip, particularly remarkable on the hackles of the cock and the wing-coverts, and also on the darker feathers of the breast.

in Occurrences 2%

The hens of both should have the body clearly pencilled across with several bars of black, and the hackle in both, sexes should be perfectly free from dark marks.

from Occurrences 1%

You're a cock of a different hackle from old Heale, then.

Which preposition to use with  hackles